Hi James! Great question.
This is something that widely varies, and depends on where you purchased the ticket, the payment method, and also the TOC’s (Train Operating Company) machine that you are using.
So for example, if you purchased a ticket on Trainline using PayPal or Apple Pay for example, then it will allow any valid debit or credit card to collect the ticket, this is because with both of those payment methods there is no way for Trainline to confirm the card number used.
It’s exactly the same if you used any type of loyalty points (RedSpottedHanky for example), either as full or part payment, then it should accept any card to collect.
In regards to payments made by card directly with the merchant, like stated above it does vary depending on the retailer and the ticket machine you are using. Even with the ticket office, I’ve seen quite a large amount of ticket collections only require a signature and not the card used. However I am especially recently seeing an increase in the amount of bookings that require the card used when collecting.
So really the rule is, sometimes it will require the original sometimes it won’t. But unless paying with Apple Pay, PayPal, loyalty points or anything that isn’t your bank card then I wouldn’t recommend not taking your card with you when collecting, just in case.